Chimarra
Chimarra | ||||||||||||
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![]() Chimarra marginata | ||||||||||||
Научная классификация | ||||||||||||
Домен: Эукариоты Царство: Животные Подцарство: Эуметазои Без ранга: Двусторонне-симметричные Без ранга: Первичноротые Без ранга: Линяющие Без ранга: Panarthropoda Тип: Членистоногие Подтип: Трахейнодышащие Надкласс: Шестиногие Класс: Насекомые Подкласс: Крылатые насекомые Инфракласс: Новокрылые Клада: Насекомые с полным превращением Надотряд: Amphiesmenoptera Отряд: Ручейники Подотряд: Кольчатощупиковые Надсемейство: Philopotamoidea Семейство: Приречные ручейники Род: Chimarra | ||||||||||||
Международное научное название | ||||||||||||
Chimarra Stephens, 1829 | ||||||||||||
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Chimarra (лат.) — род ручейников семейства Philopotamidae из подотряда Annulipalpia. Более 950 видов[1].
Распространение
Повсеместно. Род распространен на всех основных континентах, кроме Антарктиды, и делится на четыре подрода, три из которых эндемичны для Нового Света. Четвертый и номинальный подрод Chimarra встречается по всему Старому Свету, включая Африку, Азию и Австралийский регион, включая многие острова Тихого океана, а также имеет значительную радиацию в Новом Свете как в Северной, так и в Южной Америке, за исключением чилийского субрегиона. Наибольшее видовое разнообразие от мечено в Неотропике (две трети видов), Юго-Восточной Азии и Австралии. Типовой вид Chimarra marginata (Linnaeus) — единственный вид рода, широко распространенный в Европе, встречается также в Северной Африке[1][2][3][4][5][6].
Описание
Ручейники средних и мелких размеров, как правило менее 1 см. Размах крыльев представителей подрода Chimarrita от 2 до 7 мм, у Curgia — от 4 до 10 мм[1][7][8]. Личинка Chimarra распознается по наличию глубокой и асимметричной клипеальной вырезки. Цвет взрослых особей варьируется от чёрного и темно-коричневого до жёлтого и темно-красного. Средняя длина тела обычно составляет 3-8 мм, включая крылья. Общий диагноз имаго Chimarra — наличие формулы шпор 1, 4, 4; вторая анальная жилка задних крыльев сливается с первой анальной жилкой, образуя замкнутую ячейку[9].
Встречаются в разнообразных условиях и различных типах проточных пресных водоёмов, вплоть до водопадов. Личинки строят и живут в продолговатых мешкообразных фильтрующих сетях с очень мелкими ячейками, прикрепленных к нижней стороне камней на дне мелководных водоёмов[7].
- Личинка Chimarra aterrima
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Систематика
Крупнейший род ручейников, включающий чуть менее 6 % разнообразия всего отряда. Включён в подсемейство Chimarrinae. Включает более 950 видов. Выделяют четыре подрода. Подтверждение общей монофилии подродов, включая номинальный подрод, было обеспечено двумя независимыми молекулярными анализами[1][8][10][11].
Биогеографический анализ показывает происхождение рода в Неотропическом регионе и последующую быструю иррадиацию с распространением в Ориентальную, Палеарктическую и Австралазийскую области и, во вторую очередь, в Неарктическую область. Афротропический регион был колонизирован в результате нескольких независимых событий. Молекулярное датирование и калибровка по четырём ископаемым видам показывает, что роду Chimarra около 138 миллионов лет, и что иррадиация из Неотропического региона произошло примерно 124 миллиона лет назад[9]. Ископаемые виды из доминиканского янтаря (†C. dommeli , Wichard, 1983, †C. palaedominicana Wichard, 1983, †C. resinae Wichard, 1983, †C. weitschati Wichard, 1983) имеют возраст 16-20 млн лет[12].
- Подроды
- Chimarra (s. str.) Stephens, 1829
- Chimarra (Chimarrita) Blahnik, 1997
- Chimarra (Curgia) Walker, 1860
- Chimarra (Otarrha) Blahnik, 2002
- Виды
- C. aberrans AV Martynov, 1935
- C. absida Cartwright, 2020[3]
- C. abyssinica Banks, 1913
- C. aciculata JC Morse, 1974
- C. acinaciformis OS Flint, 1998
- C. actinifera F Schmid, 1958
- C. acula OS Flint, 1998
- C. acuta HH Ross, 1959
- C. adaluma DI Cartwright, 2002
- C. adamsae RJ Blahnik, 1998
- C. adelphe RJ Blahnik, 1998
- C. adella DG Denning, 1952
- C. adiatulla H Malicky, 1993
- C. adnama H Malicky, 1993
- C. africana G Enderlein, 1929
- C. aiyura K Korboot, 1965
- C. akana DG Gibbs, 1973
- C. akantha RJ Blahnik, 1997
- C. akarawitta F Schmid, 1958
- C. akkaorum P Chantaramongkol & H Malicky, 1989
- C. akruna DI Cartwright, 2002
- C. alata J Bueno-Soria, 1983
- C. alayoi L Botosaneanu, 1980
- C. albomaculata HJ Kolbe, 1888
- C. alcicorne H Malicky, 1995
- C. aliceae Cartwright, 2020[3]
- C. alticola Banks, 1937
- C. altmani RJ Blahnik, 1998
- C. alleni P Chantaramongkol & H Malicky, 1989
- C. amarganth H Malicky, 1989
- C. amazonia RJ Blahnik, 2002
- C. ambaja Mosely, 1939
- C. ambulans KH Barnard, 1934
- C. amica RJ Blahnik & RW Holzenthal, 1992
- C. aminadab H Malicky, 1993
- C. ammi H Malicky & W Graf, 2008
- C. anakwoswasi H Malicky, 1995
- C. aneca H Malicky & P Chantaramongkol, 1993
- C. angolensis G Marlier, 1965
- C. angularis RJ Blahnik, 2002
- C. angustipennis Banks, 1903
- C. anoaclana H Malicky, 1978
- C. antap Cartwright, 2020[3]
- C. antigua OS Flint, 1967
- C. antilliana OS Flint, 1968
- C. areli H Malicky & W Mey, 2008
- C. argax H Malicky, 1989
- C. argeia H Malicky & P Chantaramongkol, 1997
- C. argentella Ulmer, 1906
- C. argentinica Ulmer, 1909
- C. ariadne H Malicky, 1997
- C. arima RJ Blahnik, 1998
- C. ariomana H Malicky, 1993
- C. armata S Jacquemart, 1961
- C. assamensis DE Kimmins, 1957
- C. atara H Malicky & P Chantaramongkol, 1993
- C. aterrima HA Hagen, 1861
- C. atilanoi RJ Blahnik, 1998
- C. atnia H Malicky & P Chantaramongkol, 1993
- C. atripennis Banks, 1931
- C. augusta JC Morse, 1971
- C. aurantibasis OS Flint, 1998
- C. aureofusca DE Kimmins, 1957
- C. aureopunctata OS Flint, 1967
- C. auriceps HA Hagen, 1858
- C. auricoma DE Kimmins, 1957
- C. auripilis L Navas, 1933
- C. aurivittata OS Flint, 1971
- C. auronitens Ulmer, 1906
- C. australica Ulmer, 1916
- C. australis L Navas, 1923
- C. aviceps OS Flint, 1998
- C. babuyana W Mey, 1998
- C. bacillorum W Mey, 1990
- C. baculifera G Marlier, 1965
- C. banksi Ulmer, 1907
- C. barinas RJ Blahnik, 2002
- C. barinita OS Flint, 1998
- C. barrettae Banks, 1900
- C. batukaua H Malicky, 1995
- C. beameri DG Denning, 1950
- C. beckeri OS Flint, 1998
- C. belizensis RJ Blahnik, 1998
- C. berenike H Malicky, 1998
- C. berghei G Marlier, 1951
- C. bertrandi KMF Scott, 1974
- C. bettinae G Marlier & M Marlier, 1982
- C. beylaensis FM Gibon, 1986
- C. biatec H Malicky, 1993
- C. bibaringa DI Cartwright, 2002
- C. bicolor L Navas, 1932
- C. bicoloroides OS Flint, 1967
- C. bicornis Cartwright, 2020[3]
- C. bicuspidalis CH Sun, 1998
- C. bicuspidis Cartwright, 2020[3]
- C. bidens Ulmer, 1909
- C. bidentata RJ Blahnik, 1998
- C. bifida Cartwright, 2020[3]
- C. bimbltona H Malicky, 1979
- C. bintang Cartwright, 2020[3]
- C. biramosa DE Kimmins, 1957
- C. bisectilis OS Flint, 1998
- C. bispinosa DG Gibbs, 1973
- C. biungulata DE Kimmins, 1964
- C. blepharophera OS Flint, 1998
- C. boraceia OS Flint, 1998
- C. braconoides Walker, 1860
- C. brasiliana Ulmer, 1905
- C. briseis H Malicky, 1998
- C. bungoona DI Cartwright, 2002
- C. burmana DE Kimmins, 1957
- C. burmeisteri OS Flint, 1998
- C. butleri DG Denning, 1962
- C. caboverdensis O Nybom, 1960
- C. cachina Mosely, 1942
- C. calawiti W Mey, 1995
- C. calundoensis G Marlier, 1965
- C. callasae FM Gibon, 1982
- C. camella RJ Blahnik, 1997
- C. camerunensis G Marlier, 1980
- C. camposae OS Flint, 1998
- C. camura RJ Blahnik, 1997
- C. canoaba OS Flint, 1998
- C. cara Mosely, 1936
- C. caribea OS Flint, 1968
- C. carolae OS Flint, 1998
- C. cascada RJ Blahnik, 1998
- C. cavata Cartwright, 2020[3]
- C. cenizae W Mey, 2003
- C. centralis HH Ross, 1959
- C. centrispina OS Flint, 1998
- C. cereris KH Barnard, 1934
- C. cervicornis W Mey, 2003
- C. ceylanica DE Kimmins, 1957
- C. cipoensis OS Flint, 1998
- C. circularis HA Hagen, 1859
- C. cirrifera OS Flint, 1998
- C. clara Mosely, 1939
- C. claviloba OS Flint, 1974
- C. cogestla H Malicky, 1993
- C. cognata DE Kimmins, 1957
- C. coheni RJ Blahnik, 1998
- C. colmillo RJ Blahnik & RW Holzenthal, 1992
- C. coma H Malicky & P Chantaramongkol, 1993
- C. concava DE Kimmins, 1957
- C. concolor Ulmer, 1905
- C. confusa Ulmer, 1907
- C. conica OS Flint, 1983
- C. consimilis AV Martynov, 1912
- C. cornuta HH Ross, 1959
- C. costaricensis OS Flint, 1998
- C. creagra OS Flint, 1981
- C. crena J Bueno-Soria, 1983
- C. crenobia W Mey, 1995
- C. crepidata DE Kimmins, 1957
- C. cressae RJ Blahnik, 1998
- C. cristata Cartwright, 2020[3]
- C. crocifera JC Morse, 1974
- C. cubanorum L Botosaneanu, 1980
- C. cultellata OS Flint, 1983
- C. cumata H Malicky & P Chantaramongkol, 1993
- C. curfmani HH Ross, 1959
- C. cyclopica DE Kimmins, 1962
- C. cheesmanae DE Kimmins, 1962
- C. chela RJ Blahnik, 1997
- C. chiangmaiensis P Chantaramongkol & H Malicky, 1989
- C. chicapa G Marlier, 1965
- C. chimalapa Bueno-Soria, Santiago-Fragoso, & Barba-Alvarez, 2001
- C. chocoensis RJ Blahnik, 1998
- C. chrysosoma OS Flint, 1998
- C. damma Cartwright, 2020[3]
- C. danaokana W Mey, 1998
- C. darlingtoni RJ Blahnik, 2002
- C. decimlobata OS Flint, 1991
- C. deksamensis H Malicky, 1999
- C. demeter H Malicky, 2000
- C. denticulata Cartwright, 2020[3]
- C. dentosa HH Ross, 1948
- C. derogata A Neboiss, 2002
- C. devva H Malicky & P Chantaramongkol, 1993
- C. diakis OS Flint, 1971
- C. diannae OS Flint & JL Sykora, 1993
- C. diaphana SK Ghosh & M Chaudhury, 1999
- C. diaphora RJ Blahnik, 2002
- C. didyma OS Flint, 1998
- C. digitata AV Martynov, 1935
- C. dioni FM Gibon, 1986
- C. dirke H Malicky & P Thamsenanupap, 2000
- C. discolor DE Kimmins, 1957
- C. distermina OS Flint, 1998
- C. divergena DG Gibbs, 1973
- C. dolabrifera OS Flint & L Reyes-Arrunategui, 1991
- C. dominicana OS Flint, 1968
- C. dommeli W Wichard, 1983
- C. donamariae DG Denning & JL Sykora, 1968
- C. duckworthi OS Flint, 1967
- C. dudosa RJ Blahnik, 1998
- C. dulitensis DE Kimmins, 1955
- C. dybowskina L Navas, 1931
- C. eccaio H Malicky, 1993
- C. ediana Cartwright, 2020[3]
- C. elga Mosely, 1939
- C. elia HH Ross, 1944
- C. elviomar H Malicky, 1993
- C. embia HH Ross, 1959
- C. emima HH Ross, 1959
- C. ensifera OS Flint, 1998
- C. erecta Cartwright, 2020[3]
- C. erectiloba OS Flint, 1998
- C. espelandae Cartwright, 2020[3]
- C. espinosa RJ Blahnik, 1998
- C. evoluta DE Kimmins, 1957
- C. exapia H Malicky & P Chantaramongkol, 1993
- C. excavata DE Kimmins, 1957
- C. exilis C Sun, 2007
- C. falcata DE Kimmins, 1962
- C. falcifera S Jacquemart, 1966
- C. falculata PK Lago & SC Harris, 1987
- C. fallax Ulmer, 1912
- C. fansipangensis W Mey, 1998
- C. fenestrata DE Kimmins, 1964
- C. feria HH Ross, 1941
- C. fernandezi OS Flint, 1981
- C. feuerborni Ulmer, 1951
- C. fimbriata OS Flint, 1974
- C. fittkaui OS Flint, 1971
- C. flaviventris DE Kimmins, 1957
- C. flinti J Bueno-Soria, 1985
- C. florida HH Ross, 1944
- C. fluctuata C Sun, 2007
- C. foliata DE Kimmins, 1959
- C. forcipata RJ Blahnik, 1997
- C. formosa L Botosaneanu & R de Vos, 2006
- C. formosana Ulmer, 1915
- C. froehlichi OS Flint, 1998
- C. fulmeki Ulmer, 1951
- C. furcata S Jacquemart, 1961
- C. furti W Mey, 1998
- C. fusca DE Kimmins, 1957
- C. fuscipes DE Kimmins, 1958
- C. garciai L Botosaneanu, 1980
- C. gemmal H Malicky, 1989
- C. georgensis KH Barnard, 1934
- C. geranoides OS Flint, 1998
- C. gether H Malicky, 2009
- C. gibba RJ Blahnik, 1998
- C. gigama H Malicky, 1989
- C. gilvimacula OS Flint, 1998
- C. glaubrechti W Mey, 2006
- C. godagama F Schmid, 1958
- C. gondela OS Flint, 1974
- C. goroca JL Sykora, 1967
- C. gressitti JL Sykora, 1967
- C. guapa L Botosaneanu, 1977
- C. guatemalensis RJ Blahnik, 1998
- C. guentheri W Mey, 2006
- C. gunungkawi H Malicky, 1995
- C. guyanensis OS Flint, 1998
- C. haesitationis L Botosaneanu, 1994
- C. haimuoi H Malicky, 1995
- C. haimuoiba H Malicky, 1995
- C. haimuoibon H Malicky, 1995
- C. haimuoihai H Malicky, 1995
- C. haimuoimot H Malicky, 1995
- C. haimuoinam H Malicky, 1995
- C. hairouna L Botosaneanu, 1990
- C. hamularis C Sun, 1997
- C. harpes Cartwright, 2020[3]
- C. heliaca W Mey, 1998
- C. heligma RJ Blahnik, 1997
- C. helios H Malicky & P Chantaramongkol, 2003
- C. henryi DE Kimmins, 1957
- C. heppneri RJ Blahnik, 1997
- C. hezron H Malicky, 1993
- C. hienghene H Malicky, 1981
- C. hoangliensis W Mey, 2005
- C. holzenthali PK Lago & SC Harris, 1987
- C. hoogstraali HH Ross, 1956
- C. horok H Malicky, 1989
- C. houvichka F Schmid, 1960
- C. htinorum P Chantaramongkol & H Malicky, 1989
- C. huonana Cartwright, 2020[3]
- C. hyoeides OS Flint, 1983
- C. igvarvaria SI Melnitsky, 2005
- C. immaculata Ulmer, 1911
- C. incipiens RJ Blahnik, 2002
- C. indigota Mosely, 1941
- C. inflata RJ Blahnik, 1998
- C. ino G Marlier, 1981
- C. intermedia S Jacquemart, 1961
- C. intexta Mosely, 1931
- C. inthanonensis P Chantaramongkol & H Malicky, 1989
- C. irwini OS Flint, 1998
- C. ismayi Cartwright, 2020[3]
- C. izabala RJ Blahnik, 1998
- C. jabal H Malicky & P Chantaramongkol, 2009
- C. jacobsoni Ulmer, 1951
- C. jamaicensis OS Flint, 1968
- C. janzeni RJ Blahnik & RW Holzenthal, 1992
- C. jari Cartwright, 2020[3]
- C. jaroschi H Malicky, 1994
- C. jemima RJ Blahnik & RW Holzenthal, 1992
- C. jiraprapa P Chantaramongkol & H Malicky, 1986
- C. jisipu H Malicky, 1989
- C. johansoni Cartwright, 2020[3]
- C. joliveti S Jacquemart, 1979
- C. jugescens OS Flint, 1998
- C. juliae OS Flint, 1998
- C. kabashana G Marlier, 1943
- C. kailishchandrai H Malicky, 1997
- C. kaiya DI Cartwright, 2002
- C. karakara DI Cartwright, 2002
- C. karenorum P Chantaramongkol & H Malicky, 1989
- C. kenyana Ulmer, 1931
- C. kewabi Cartwright, 2020[3]
- C. kewarra DI Cartwright, 2002
- C. khamuorum P Chantaramongkol & H Malicky, 1989
- C. khasia DE Kimmins, 1957
- C. koki L Botosaneanu, 1996
- C. kokodana DE Kimmins, 1962
- C. kontilos RJ Blahnik, 1997
- C. koualeensis KA Johanson & N Mary, 2009[13]
- C. krugeri S Jacquemart, 1963
- C. kuala J Olah, 1993
- C. kuka Cartwright, 2020[3]
- C. kumaonensis AV Martynov, 1935
- C. kwansiensis C Hwang, 1957
- C. lacroixi L Navas, 1921
- C. laguna HH Ross, 1951
- C. lahuorum P Chantaramongkol & H Malicky, 1989
- C. langleyae RJ Blahnik, 1998
- C. lankana DE Kimmins, 1957
- C. lannaensis P Chantaramongkol & H Malicky, 1989
- C. larapinta DI Cartwright, 2002
- C. lata RJ Blahnik & RW Holzenthal, 1992
- C. latiloba W Mey, 2003
- C. lavuaorum P Chantaramongkol & H Malicky, 1989
- C. lejea Mosely, 1948
- C. leopoldi S Jacquemart, 1981
- C. leta Mosely, 1936
- C. leucophlebia L Navas, 1932
- C. lewisi DE Kimmins, 1957
- C. lichiuensis LP Hsu & CS Chen, 1996
- C. limon RJ Blahnik, 1998
- C. lissuorum P Chantaramongkol & H Malicky, 1989
- C. litugena H Malicky & P Chantaramongkol, 1993
- C. litussa H Malicky & P Chantaramongkol, 1993
- C. lobata OS Flint, 1967
- C. locolo DI Cartwright, 2002
- C. lojaensis OS Flint, 1998
- C. longispina C Sun, 2007
- C. longistylis S Jacquemart & B Statzner, 1981
- C. longiterga RJ Blahnik & RW Holzenthal, 1992
- C. longpela DI Cartwright, 2001
- C. lorengau H Malicky, 1994
- C. loriana L Navas, 1933
- C. lotta H Malicky, 1993
- C. lucretia H Malicky & P Thamsenanupap, 2005
- C. lufirae S Jacquemart, 1961
- C. lukawaei S Jacquemart, 1961
- C. luminaris DI Cartwright, 2002
- C. lupialae S Jacquemart, 1961
- C. luzonica Banks, 1913
- C. luzoquezonensis W Mey, 2003
- C. lwirona B Statzner, 1976
- C. macara OS Flint, 1998
- C. machaerophora OS Flint, 1968
- C. magsaysayensis W Mey, 2003
- C. majuscula RJ Blahnik, 1997
- C. malaisei DE Kimmins, 1957
- C. maldonadoi OS Flint, 1964
- C. manni Banks, 1924
- C. maoi CH Sun & H Malicky, 2002
- C. margaritae OS Flint, 1991
- C. marginata C Linnaeus, 1767
- C. maritza OS Flint, 1998
- C. mars H Malicky, 2007
- C. massana H Malicky, 1994
- C. matura H Malicky & P Chantaramongkol, 1993
- C. mauritania S Jacquemart, 1960
- C. mayottensis KA Johanson & N Mary, 2009[13]
- C. medioloba OS Flint, 1971
- C. megara H Malicky, 2007
- C. meorum P Chantaramongkol & H Malicky, 1989
- C. merengue RJ Blahnik, 1997
- C. mexicana Banks, 1900
- C. minca OS Flint, 1998
- C. mindanensis W Mey, 1998
- C. minga OS Flint, 1998
- C. minima Ulmer, 1907
- C. minuta AV Martynov, 1935
- C. mitis HA Hagen, 1858
- C. mlabriorum P Chantaramongkol & H Malicky, 1989
- C. moesta Banks, 1924
- C. moira H Malicky & T Prommi, 2004
- C. momma H Malicky & P Chantaramongkol, 1993
- C. mommaides W Mey, 1998
- C. mongelutonga H Malicky, 1979
- C. monorum P Chantaramongkol & H Malicky, 1989
- C. montana DE Kimmins, 1955
- C. monticola DE Kimmins, 1953
- C. morio H Burmeister, 1839
- C. moselyi DG Denning, 1948
- C. mouldsi DI Cartwright, 2002
- C. munozi RJ Blahnik & RW Holzenthal, 1992
- C. muoibay H Malicky, 1995
- C. muoichin H Malicky, 1995
- C. muoitam H Malicky, 1995
- C. mushuvae G Marlier, 1951
- C. mussaua H Malicky, 1994
- C. mycterophora OS Flint, 1998
- C. nabilla DI Cartwright, 2002
- C. nahesson H Malicky & P Chantaramongkol, 1993
- C. nasuta OS Flint, 1998
- C. natalicia DI Cartwright, 2002
- C. neblina RJ Blahnik, 1997
- C. neboissi DI Cartwright, 2002
- C. nemet H Malicky, 1993
- C. neofimbriata OS Flint, 1974
- C. nepalensis DE Kimmins, 1964
- C. nervosa Brauer, 1867
- C. nigra DE Kimmins, 1964
- C. nigrella W Mey, 1995
- C. nigrorosea F Schmid, 1960
- C. noebia H Malicky & P Chantaramongkol, 1993
- C. nonna H Malicky, 1993
- C. nunenada SI Melnitsky, 2005
- C. oaxaca RJ Blahnik, 1998
- C. obscura Walker, 1852
- C. obscurella Banks, 1924
- C. odonta RJ Blahnik, 2002
- C. oinone H Malicky, 2007
- C. okuihorum W Mey, 1998
- C. onima OS Flint, 1991
- C. opaca W Mey, 1998
- C. ophiognatha W Mey, 1998
- C. oreithyia H Malicky, 2007
- C. ortiziana OS Flint, 1967
- C. orumbera DI Cartwright, 2002
- C. otuzcoensis OS Flint & L Reyes-Arrunategui, 1991
- C. ovalis HH Ross, 1959
- C. pablito OS Flint, 1998
- C. palaedominicana W Wichard, 1983
- C. palaenova W Wichard, 2007
- C. palawana H Malicky, 1994
- C. panguna DI Cartwright, 2001
- C. papuana DE Kimmins, 1962
- C. paracreagra RJ Blahnik, 1998
- C. parana OS Flint, 1972
- C. paraortiziana RJ Blahnik & RW Holzenthal, 1992
- C. parasocia PK Lago & SC Harris, 1987
- C. parene RJ Blahnik, 2002
- C. paria OS Flint, 1998
- C. particeps RJ Blahnik, 2002
- C. pasiphae H Malicky, 2007
- C. pataplan H Malicky, 1989
- C. patosa HH Ross, 1956
- C. paucispina AP Moreira Santos & JL Nessimian, 2009
- C. pedalis Banks, 1931
- C. peineta RJ Blahnik & RW Holzenthal, 1992
- C. pelaezi J Bueno-Soria, 1985
- C. persimilis Banks, 1920
- C. peruana RJ Blahnik, 2002
- C. peruviana OS Flint, 1998
- C. petersorum OS Flint, 1998
- C. petri DG Gibbs, 1973
- C. petricola OS Flint, 1998
- C. peytoni OS Flint, 1998
- C. philipponi FM Gibon, 1986
- C. phlegyas H Malicky, 2008
- C. phthanorossi RJ Blahnik, 2002
- C. phuwa H Malicky & P Chantaramongkol, 2009
- C. picea L Navas, 1924
- C. piliferosa OS Flint, 1998
- C. pilosella L Navas, 1932
- C. pillara DI Cartwright, 2002
- C. pinga DI Cartwright, 2001
- C. pipake H Malicky & P Chantaramongkol, 1993
- C. piraya OS Flint, 1983
- C. pita DI Cartwright, 2002
- C. platyrhina OS Flint, 1981
- C. plaumanni OS Flint, 1983
- C. podarge H Malicky & P Thamsenanupap, 2007
- C. polyneikes H Malicky, 2008
- C. pollex RJ Blahnik & RW Holzenthal, 1992
- C. pondoensis KH Barnard, 1941
- C. pontos H Malicky, 2007
- C. poolei OS Flint, 1981
- C. potamophila W Mey, 1995
- C. potosi RJ Blahnik, 1998
- C. primula DG Denning, 1950
- C. prisna P Chantaramongkol & H Malicky, 1986
- C. prodhoni FM Gibon, 1985
- C. prokrustes H Malicky, 2008
- C. prolata RJ Blahnik, 1997
- C. protuberans RJ Blahnik, 1998
- C. puertoricensis OS Flint, 1964
- C. pulchra HA Hagen, 1861
- C. pulla L Navas, 1932
- C. pumila Banks, 1920
- C. purisca OS Flint, 1998
- C. pusilla RJ Blahnik, 1997
- C. puya OS Flint, 1998
- C. pylaea DG Denning, 1941
- C. quadratiterga RJ Blahnik, 1998
- C. quadridigitata Yang, Sun & Yang, 2001
- C. quadrifurcata L Botosaneanu, 1994
- C. quadrispinosa S Jacquemart & B Statzner, 1981
- C. quaternaria OS Flint, 1971
- C. quina OS Flint, 1998
- C. quitacalzon RJ Blahnik, 1998
- C. rafita RJ Blahnik, 1998
- C. ram H Malicky, 1993
- C. rama H Malicky & P Chantaramongkol, 1993
- C. ramakien H Malicky & P Chantaramongkol, 1993
- C. ranuka DI Cartwright, 2002
- C. ravanna H Malicky & P Chantaramongkol, 1993
- C. reasilvia H Malicky & T Prommi, 2006
- C. recta Ulmer, 1930
- C. redonda RJ Blahnik, 2002
- C. regu H Malicky & P Chantaramongkol, 2009
- C. resinae W Wichard, 1983
- C. retrorsa OS Flint, 1974
- C. reyangensis SK Ghosh & M Chaudhury, 1999
- C. rhamphodes RJ Blahnik, 1998
- C. rhodesi DE Kimmins, 1957
- C. ridleyi DG Denning, 1941
- C. robynsi S Jacquemart, 1967
- C. rosalesi OS Flint, 1981
- C. rossi J Bueno-Soria, 1985
- C. ruficeps Ulmer, 1913
- C. sabrona DE Kimmins, 1962
- C. sadayu H Malicky, 1993
- C. saganeitina L Navas, 1932
- C. sandhamma F Schmid, 1958
- C. sarophora OS Flint, 1998
- C. sassandrae FM Gibon, 1982
- C. saudia H Malicky, 1986
- C. scopula OS Flint, 1974
- C. scopulifera DE Kimmins, 1957
- C. scopuloides OS Flint, 1974
- C. schiza HH Ross, 1959
- C. schmidi DE Kimmins, 1962
- C. schwendingeri P Chantaramongkol & H Malicky, 1989
- C. securigera OS Flint, 1998
- C. sedlaceki JL Sykora, 1967
- C. segmentipennis C Hwang, 1957
- C. semiramis H Malicky, 2007
- C. sensillata OS Flint, 1981
- C. senticosa CH Sun & H Malicky, 2002
- C. septemlobata OS Flint, 1991
- C. septifera OS Flint, 1974
- C. sepulchralis HA Hagen, 1858
- C. setosa HH Ross, 1959
- C. shanorum P Chantaramongkol & H Malicky, 1989
- C. shaowuensis C Hwang, 1957
- C. shiva H Malicky & P Chantaramongkol, 1993
- C. siami S Jacquemart, 1979
- C. signata Banks, 1936
- C. simplaria W Mey, 2003
- C. simpliciforma OS Flint, 1971
- C. sinuata C Hwang, 1957
- C. sinuosa DE Kimmins, 1962
- C. sita H Malicky & P Chantaramongkol, 1993
- C. skaidan H Malicky, 1989
- C. skiborskii W Mey, 1995
- C. socia HA Hagen, 1861
- C. solisi RJ Blahnik & RW Holzenthal, 1992
- C. somereni G Marlier, 1951
- C. spangleri J Bueno-Soria, 1985
- C. spatulata HH Ross, 1959
- C. spinifera DE Kimmins, 1957
- C. spinulifera OS Flint, 1968
- C. spitzeri H Malicky, 1994
- C. stclairae DI Cartwright, 2002
- C. straminea OS Flint, 1998
- C. strongyla RJ Blahnik, 1998
- C. suadulla H Malicky & P Chantaramongkol, 1993
- C. succini W Wichard, 1983
- C. supanna H Malicky, 1993
- C. suryasena F Schmid, 1960
- C. suthepensis P Chantaramongkol & H Malicky, 1989
- C. sylvestris FM Gibon, 1985
- C. sythoffi Ulmer, 1951
- C. szunyoghyi J Olah, 1986
- C. tachuela RJ Blahnik, 2002
- C. tagaijana W Mey, 2003
- C. tagalica Banks, 1937
- C. talos H Malicky, 2007
- C. tallawalla DI Cartwright, 2002
- C. tamba OS Flint, 1998
- C. tamsi Mosely, 1936
- C. tapanti RJ Blahnik, 1998
- C. tawitawi H Malicky, 1994
- C. telihigola F Schmid, 1958
- C. teresae OS Flint, 1998
- C. terramater H Malicky, 2008
- C. texana Banks, 1920
- C. thaiorum P Chantaramongkol & H Malicky, 1989
- C. thaumas H Malicky, 2008
- C. thienemanni Ulmer, 1951
- C. thyestes H Malicky, 2008
- C. tibialis L Navas, 1922
- C. tityos H Malicky, 2008
- C. toga H Malicky & P Chantaramongkol, 1993
- C. togoana Ulmer, 1907
- C. toliana Cartwright, 2020[3]
- C. tortuosa RJ Blahnik, 1997
- C. toubaensis FM Gibon, 1985
- C. travei S Jacquemart, 1963
- C. triangularis DE Kimmins, 1963
- C. triangulata LP Hsu & CS Chen, 1996
- C. trigona Cartwright, 2020[3]
- C. trispina S Jacquemart, 1961
- C. truncatiloba OS Flint, 1974
- C. tsudai HH Ross, 1956
- C. tucuna OS Flint, 1998
- C. uara OS Flint, 1971
- C. ulmeri DE Kimmins, 1962
- C. uncata JC Morse, 1974
- C. uncula W Mey, 1998
- C. upia H Malicky, 1993
- C. uppita H Malicky & P Chantaramongkol, 1993
- C. uranka Mosely, 1953
- C. usai H Malicky & P Chantaramongkol, 2009
- C. uschtu H Malicky, 1989
- C. usitatissima OS Flint, 1971
- C. utahensis HH Ross, 1938
- C. utra RJ Blahnik, 1998
- C. uvana DE Kimmins, 1957
- C. uvirana G Marlier, 1951
- C. vasoudeva F Schmid, 1960
- C. vesta H Malicky, 2007
- C. vibena H Malicky & P Chantaramongkol, 1993
- C. villalobosi J Bueno-Soria, 1985
- C. virgencita RJ Blahnik & RW Holzenthal, 1992
- C. waensis FM Gibon, 1985
- C. weitschati W Wichard, 1983
- C. wiharawela F Schmid, 1958
- C. wilcuma RJ Blahnik, 1998
- C. wilsoni OS Flint, 1967
- C. woldai RJ Blahnik, 1998
- C. wooroonooran DI Cartwright, 2002
- C. wushikangensis LP Hsu & CS Chen, 1996
- C. xenillion A Neboiss, 1986
- C. xingu RJ Blahnik, 1997
- C. xus RJ Blahnik, 1998
- C. yaloma H Malicky, 1994
- C. yandala DI Cartwright, 2002
- C. yanura RJ Blahnik & RW Holzenthal, 1992
- C. yaorum P Chantaramongkol & H Malicky, 1989
- C. yaoshanensis C Hwang, 1957
- C. yoolumba DI Cartwright, 2002
- C. ypsilon OS Flint, 1983
- C. yskal H Malicky, 1989
- C. yulae DI Cartwright, 2001
- C. zagrosensis P Chvojka, 1996
- C. zamora RJ Blahnik, 1998
- C. zilla H Malicky & P Chantaramongkol, 2009
- C. zoria Mosely, 1939
Примечания
- ↑ 1 2 3 4 Blahnik R., Andersen T. New species of the genus Chimarra Stephens from Africa (Trichoptera, Philopotamidae) and characterization of the African groups and subgroups of the genus. In: Pauls SU, Thomson R, Rázuri-Gonzales E (Eds) Special Issue in Honor of Ralph W. Holzenthal for a Lifelong Contribution to Trichoptera Systematics. // ZooKeys. — 2022. — Т. 1111. — С. 43—198. — ISSN 1869-4802 1867-6294, 1869-4802. — doi:10.3897/zookeys.1111.77586.
- ↑ Cartwright David I. The Australian species of Chimarra Stephens (Trichoptera: Philopotamidae) // Memoirs of the Museum of Victoria. — 2002. — Vol. 59(2). — С. 393—437. — doi:10.24199/j.mmv.2002.59.8.
- ↑ 1 2 3 4 5 6 7 8 9 10 11 12 13 14 15 16 17 18 19 20 21 22 23 24 Cartwright D. I. A review of the New Guinea species of Chimarra Stephens (Trichoptera: Philopotamidae) // Memoirs of the Museum of Victoria[англ.]. — Museums Victoria[англ.], 2020. — Vol. 79. — С. 1—49. — ISSN 1447-2554. — doi:10.24199/j.mmv.2020.79.01.
- ↑ Johanson K. A., Espeland M. Description of new Chimarra (Trichoptera: Philopotamidae) species from the Solomon Islands // Zootaxa. — 2010. — Vol. 2638(1). — С. 25—43. — doi:10.11646/zootaxa.2638.1.2.
- ↑ Johanson K. A., Wells A., Malm T., Espeland M. The Trichoptera of Vanuatu // Deutsche Entomologische Zeitschrift. — 2011. — Vol. 58(2). — С. 279—320. — doi:10.1002/mmnd.201100031.
- ↑ Johanson K. A., Oláh J. Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species // Zootaxa. — 2012. — Vol. 3354(1). — С. 1—58. — doi:10.11646/zootaxa.3354.1.1.
- ↑ 1 2 Blahnik R. J. Systematics of Chimarrita, a new subgenus of Chimarra (Trichoptera: Philopotamidae) (англ.) // Systematic Entomology : Журнал. — 1997. — Vol. 22 (3). — P. 199—243. — doi:10.1046/j.1365-3113.1997.d01-39.x.
- ↑ 1 2 Flint Jr O.S. Studies of Neotropical caddisflies, LIII: a taxonomic revision of the subgenus Curgia of the genus Chimarra (Trichoptera: Philopotamidae) (англ.) // Smithsonian Contributions to Zoology : Журнал. — 1998. — Vol. 594. — P. 1—131. — doi:10.5479/si.00810282.594.
- ↑ 1 2 Wahlberg E., Johanson K.A. The age, ancestral distribution and radiation of Chimarra (Trichoptera: Philopotamidae) using molecular methods (англ.) // Molecular Phylogenetics and Evolution : Журнал. — 2014. — Vol. 79. — P. 433—442. — doi:10.1016/j.ympev.2014.06.023.
- ↑ Kjer KM, Xhou X, Frandsen PB, Thomas JA, Blahnik RJ. Moving toward species-level phylogeny using ribosomal DNA and COI barcodes: an example from the diverse caddisfly genus Chimarra (Trichoptera: Philopotamidae) (англ.) // Arthropod Systematics & Phylogeny : Журнал. — 2014. — Vol. 72, no. 3. — P. 345—354.
- ↑ Blahnik Roger J. Systematics of Otarrha, a new Neotropical subgenus of Chimarra (Trichoptera: Philopotamidae) (англ.) // Systematic Entomology : Журнал. — 2002. — Vol. 27, no. 1. — P. 65—130. — doi:10.1046/j.0307-6970.2001.00166.x.
- ↑ Wichard W. Köcherfliegen des Dominikanischen Bernsteins II. Fossile Arten der Gattung Chimarra (Trichoptera, Philopotamidae) (нем.) // Mitt. Münchner Entomol. Gesell. : Журнал. — 1983. — Vol. 72. — P. 137—145.
- ↑ 1 2 Johanson, Kjell Arne, Mary, Nathalie (2009): Description of three new caddisfly species from Mayotte Island, Comoros Archipelago (Insecta: Trichoptera). Zootaxa 2089: 1-9, DOI: 10.5281/zenodo.187447
Литература
- Blahnik R. J. Systematics of Chimarrita, a new subgenus of Chimarra (Trichoptera: Philopotamidae) (англ.) // Systematic Entomology : Журнал. — 1997. — Vol. 22 (3). — P. 199—243. — doi:10.1046/j.1365-3113.1997.d01-39.x.
- Blahnik R. J. Systematics of Otarrha, a new Neotropical subgenus of Chimarra (Trichoptera: Philopotamidae) (англ.) // Systematic Entomology : Журнал. — 2002. — Vol. 27 (1). — P. 65—130. — doi:10.1046/j.0307-6970.2001.00166.x.
- Blahnik Roger, Ralph Holzenthal. New Neotropical species of Chimarra (Trichoptera, Philopotamidae), ZooKeys, 10.3897/zookeys.184.2911, 184, (1-33), (2012).
- Flint Jr OS. (1998). Studies of Neotropical caddisflies, LIII: a taxonomic revision of the subgenus Curgia of the genus Chimarra (Trichoptera: Philopotamidae). Smithsonian Contributions to Zoology 594: 1-131. https://doi.org/10.5479/si.00810282.594
- Johanson K. A., J. Oláh. Revision of the Fijian Chimarra (Trichoptera, Philopotamidae) with description of 24 new species (англ.) // Zootaxa : Журнал. — Auckland, New Zealand: Magnolia Press, 2012. — Vol. 3354. — P. 1—58. — ISSN 1175-5326. — doi:10.11646/zootaxa.3354.1.1.
- François-Marie Gibon. The Chimarra minima group in West Africa and Madagascar (Trichoptera, Philopotamidae), Zoosystema, 10.5252/z2015n2a3, 37, 2, (333—350), (2015).
- Albane Vilarino, Adolfo Calor. New species and records of Chimarra (Trichoptera, Philopotamidae) from Northeastern Brazil, and an updated key to subgenus Chimarra (Chimarrita), ZooKeys, 10.3897/zookeys.491.8553, 491, (119—142), (2015).
Ссылки
- Philopotamidae — Tree of Life
- Trichoptera World Checklist