Phyllophaga

Phyllophaga

Nomenklatur
Phyllophaga Harris, 1827
Synonymi
Tostegoptera Blanchard, 1851, Trichestes Blanchard, 1851, Eugastra LeConte, 1854, Endrosa LeConte, 1856, Gynnis LeConte, 1856, Chirodines Bates, 1888, Abcrana Saylor, 1942, Clemora Saylor, 1942, Cnemarachis Saylor, 1942
Hører til
skarabider, Polyphaga, biller
Økologi
Antall arter:		ca. 840
Habitat:		i jord, gressmarker
Utbredelse:		Amerika
Inndelt i

Phyllophaga er en meget artsrik slekt av biller som hører til gruppen oldenborrer (Melolonthinae) i den store gruppen skarabider (Scarabaeoidea).

Utseende

Små til ganske store (12-35 millimeter), sylindriske, brunlige oldenborrer. Kroppen er forholdsvis tynnskallet og mangler skjell og påfallende hårkledning på oversiden.

Levevis

Larvene lever i jorda som vanlig for oldenborrer. Noen av artene kan bli svært tallrike og gjøre skade i landbruket. De voksne billene gnager på mange slags planter.

Utbredelse

Slekten er utbredt i Amerika, fra Argentina i sør til Canada i nord, inkludert øyene i Karibia. Rundt halvparten av artene lever i Nord-Amerika.

Systematisk inndeling

Treliste

Ordenen biller, Coleoptera
- Underordenen Polyphaga
  - Overfamilien skarabider, Scarabaeoidea
    - Familien skarabider, Scarabaeidae Latreille, 1806
      - Underfamilien oldenborrer, Melolonthinae Samouelle, 1819
        Stammen Rhizotrogini Burmeister, 1855
        Slekten Phyllophaga Harris, 1827

Liste over artene

Phyllophaga abcea Saylor, 1940
Phyllophaga abdominalis (Moser, 1921)
Phyllophaga abudantuni Chalumeau & Gruner, 1976
Phyllophaga acacoyahuana Morón & Blas, 2006
Phyllophaga acapulca Saylor, 1943
Phyllophaga acatlanensis Morón & Aragon-García, 2012
Phyllophaga aceitillar Woodruff, 2005
Phyllophaga acinosa (Arrow, 1920)
Phyllophaga acunai Chapin, 1937
Phyllophaga adjuntas Saylor, 1940
Phyllophaga aegrotus
Phyllophaga aemula (Horn, 1887)
Phyllophaga aenea (Moser, 1921)
Phyllophaga aeneotincta Chapin, 1932
Phyllophaga aequalis (LeConte, 1853)
Phyllophaga aequatorialis (Moser, 1921)
Phyllophaga aeruginosa (Burmeister, 1855)
Phyllophaga affabilis (Horn, 1887)
Phyllophaga afflicta (Blanchard, 1850)
Phyllophaga aguadita Saylor, 1942
Phyllophaga ahlbrandti García-Vidál, 1988
Phyllophaga alayoi García-Vidál, 1978
Phyllophaga albina (Burmeister, 1855)
Phyllophaga alcoa Woodruff, 2005
Phyllophaga aliada Sanderson, 1951
Phyllophaga aliciae Morón & Riva-Cervantes, 2005
Phyllophaga alquizara Chapin, 1932
Phyllophaga alvareztoroi Morón & Blas, 2006
Phyllophaga amazonica (Moser, 1918)
Phyllophaga analis (Burmeister, 1855)
Phyllophaga andersoni Smith & Paulsen, 2015
Phyllophaga androw Woodruff, 2005
Phyllophaga angulicollis (Bates, 1887)
Phyllophaga angusta (Blanchard, 1850)
Phyllophaga anolaminata (Moser, 1921)
Phyllophaga anomaloides (Bates, 1887)
Phyllophaga antennalis (Moser, 1921)
Phyllophaga antiguae (Arrow, 1920)
Phyllophaga anxia (LeConte, 1850)
Phyllophaga apicalis (Blanchard, 1850)
Phyllophaga apolinari Saylor, 1940
Phyllophaga apolinaria Saylor, 1942
Phyllophaga approxima Woodruff & Sanderson, 2005
Phyllophaga aragoni Morón, 2013
Phyllophaga arcta (Horn, 1887)
Phyllophaga ardara Saylor, 1943
Phyllophaga arkansana (Schaeffer, 1906)
Phyllophaga arribans Saylor, 1943
Phyllophaga arrowi (Saylor, 1935)
Phyllophaga assmani Saylor, 1936
Phyllophaga atra (Moser, 1918)
Phyllophaga atrata (Moser, 1918)
Phyllophaga atratoides Morón, 2003
Phyllophaga audanti Wolcott, 1928
Phyllophaga aurea Luginbill & Painter, 1941
Phyllophaga austera (Erichson, 1847)
Phyllophaga babis Saylor, 1943
Phyllophaga badbunnyi Cano, 2020
Phyllophaga bahiana (Saylor, 1935)
Phyllophaga balia (Say, 1825)
Phyllophaga balli Morón & Woodruff, 2008
Phyllophaga baneta Saylor, 1943
Phyllophaga baoruco Woodruff, 2005
Phyllophaga baracoana Chapin, 1932
Phyllophaga baraguensis Chapin, 1932
Phyllophaga barda (Horn, 1887)
Phyllophaga baroni (Bates, 1890)
Phyllophaga barrapotosiana Romero & Morón, 2017
Phyllophaga barroi Serrano & Morón, 2017
Phyllophaga barrosa Sanderson, 1951
Phyllophaga batillifer (Bates, 1887)
Phyllophaga beameri Sanderson, 1958
Phyllophaga beckeri (Moser, 1921)
Phyllophaga benexonana Morón, 2000
Phyllophaga bicavifrons Chapin, 1932
Phyllophaga bicolor (Moser, 1921)
Phyllophaga bilobatata Saylor, 1939
Phyllophaga bilyi Chalumeau, 1989
Phyllophaga bimaculata García-Vidál, 1984
Phyllophaga bimammifrons Saylor, 1940
Phyllophaga bipartita (Horn, 1887)
Phyllophaga blackwelderi Saylor, 1940
Phyllophaga blanchardi (Arrow, 1933)
Phyllophaga blanda Sanderson, 1958
Phyllophaga bobevelynorum Schnepp, 2022
Phyllophaga bolacoides (Bates, 1887)
Phyllophaga boliviensis (Blanchard, 1850)
Phyllophaga bonfils Woodruff & Sanderson, 2005
Phyllophaga boruca Morón, 2003
Phyllophaga bowditchi Saylor, 1938
Phyllophaga brama Saylor, 1943
Phyllophaga bretti Smith & Paulsen, 2015
Phyllophaga brevicornis García-Vidál, 1988
Phyllophaga brevipilosa (Moser, 1918)
Phyllophaga brownella Saylor, 1942
Phyllophaga bruneri Chapin, 1932
Phyllophaga bueta Saylor, 1940
Phyllophaga buapae Morón, 2013
Phyllophaga bucephala (Bates, 1887)
Phyllophaga caanchaki Morón, 1998
Phyllophaga cahitana Morón, 2001
Phyllophaga calai Smith & Paulsen, 2015
Phyllophaga calceata (LeConte, 1856)
Phyllophaga calderasa (Saylor, 1937)
Phyllophaga callosiventris (Moser, 1921)
Phyllophaga cambeforti Cartwright & Chalumeau, 1977
Phyllophaga campana Morón & Riley, 2005
Phyllophaga caneyensis García-Vidál, 1984
Phyllophaga canipolea Saylor, 1948
Phyllophaga canoa Sanderson, 1951
Phyllophaga canoana Morón, 2003
Phyllophaga capillata (Blanchard, 1850)
Phyllophaga caraga Saylor, 1943
Phyllophaga cardini Chapin, 1932
Phyllophaga carnegie Woodruff, 2005
Phyllophaga cartaba Sanderson, 1951
Phyllophaga castaniella (Bates, 1887)
Phyllophaga castineirasi García-Vidál, 1978
Phyllophaga catemacoana Morón, 2003
Phyllophaga caviceps (Moser, 1918)
Phyllophaga cayennensis (Moser, 1918)
Phyllophaga caymanensisSanderson, 1939
Phyllophaga cazahuata Morón, 2013
Phyllophaga cazieriana Saylor, 1938
Phyllophaga cececpana Morón, 2016
Phyllophaga centralis (Nonfried, 1894)
Phyllophaga certanca Saylor, 1943
Phyllophaga chada Saylor, 1948
Phyllophaga chalumeaui García-Vidál, 1984
Phyllophaga chamacayoca Morón, 1991
Phyllophaga chamula Morón, 1999
Phyllophaga changuena Morón, 2003
Phyllophaga chiapas Saylor, 1943
Phyllophaga chiblacana Morón, 2003
Phyllophaga chimoxtila Morón, 2003
Phyllophaga chippewa
Phyllophaga chiriquina (Bates, 1887)
Phyllophaga chlaenobiana Saylor, 1936
Phyllophaga cholana Morón, 2003
Phyllophaga chortiana Morón, 2003
Phyllophaga cinnamomea (Blanchard, 1850)
Phyllophaga citarae Morón & Neita-Moreno, 2014
Phyllophaga ciudadensis (Bates, 1887)
Phyllophaga clavijeroi Morón, 2015
Phyllophaga clemens (Horn, 1887)
Phyllophaga clypeata (Horn, 1887)
Phyllophaga cneda Saylor, 1940
Phyllophaga coahuayana Morón, 2006
Phyllophaga cocleana Morón, 2013
Phyllophaga colima Saylor, 1943
Phyllophaga colimana (Moser, 1921)
Phyllophaga collaris (Moser, 1921)
Phyllophaga collaroides Morón, Lugo-Gracía & Aragon-García, 2015
Phyllophaga columbiana (Blanchard, 1850)
Phyllophaga comaltepecana Morón, 2003
Phyllophaga cometes (Bates, 1887)
Phyllophaga complexipennis García-Vidál, 1984
Phyllophaga conformis (Blanchard, 1850)
Phyllophaga congrua (LeConte, 1856)
Phyllophaga conicariana Morón, 2006
Phyllophaga contaminata Fall, 1932
Phyllophaga coronadis Saylor, 1941
Phyllophaga corrosa (LeConte, 1856)
Phyllophaga costaricensis (Moser, 1918)
Phyllophaga costura Sanderson, 1951
Phyllophaga crassa (Burmeister, 1855)
Phyllophaga crassissima (Blanchard, 1850)
Phyllophaga crena Saylor, 1941
Phyllophaga crenaticollis (Blanchard, 1850)
Phyllophaga crenonycha Saylor, 1943
Phyllophaga crenulata (Frölich, 1792)
Phyllophaga cribrosa (LeConte, 1853)
Phyllophaga crinipennis (Bates, 1890)
Phyllophaga crinitissima Wolcott, 1924
Phyllophaga cristobala Saylor, 1942
Phyllophaga cruces Morón & Riley, 2005
Phyllophaga cubana Chapin, 1932
Phyllophaga cupuliformis Langston, 1924
Phyllophaga curialis Reinhard, 1939
Phyllophaga cuyabana (Moser, 1918)
Phyllophaga cylindrica (Burmeister, 1855)
Phyllophaga darlingtoni Sanderson, 1940
Phyllophaga dasypoda (Bates, 1887)
Phyllophaga davidsoni Woodruff, 2005
Phyllophaga davisi Langston, 1927
Phyllophaga debilis (LeConte, 1856)
Phyllophaga delata (Horn, 1887)
Phyllophaga deleri Smith & Paulsen, 2015
Phyllophaga delphinicauda Androw, 2016
Phyllophaga delplanquei Chalumeau & Gruner, 1976
Phyllophaga densata (Moser, 1918)
Phyllophaga densepunctata (Moser, 1918)
Phyllophaga dentex (Bates, 1887)
Phyllophaga denticulata (Blanchard, 1850)
Phyllophaga dieteriana Deloya & Morón, 1998
Phyllophaga diffinis (Blanchard, 1850)
Phyllophaga diminuta Evans, 2003
Phyllophaga disca Saylor, 1943
Phyllophaga discalis Chapin, 1935
Phyllophaga dispar (Burmeister, 1855)
Phyllophaga dissimilis (Chevrolat, 1865)
Phyllophaga divertens (Bates, 1887)
Phyllophaga dominicensis Cartwright & Chalumeau, 1977
Phyllophaga drakii (Kirby, 1837)
Phyllophaga dsaimana Morón, 2003
Phyllophaga dubitata García-Vidál, 1978
Phyllophaga duenas Saylor, 1941
Phyllophaga dugesiana Morón, Rivera & López, 2001
Phyllophaga dulcis (Bates, 1887)
Phyllophaga durangoana (Moser, 1921)
Phyllophaga durangosa Saylor, 1943
Phyllophaga ecostata (Horn, 1887)
Phyllophaga ecuadorae Blackwelder, 1944
Phyllophaga edrileyi Morón, 2015
Phyllophaga eladio Woodruff, 2005
Phyllophaga elegans (Nonfried, 1891)
Phyllophaga elenans Saylor, 1938
Phyllophaga elizoria (Saylor, 1937)
Phyllophaga elongata (Linell, 1896)
Phyllophaga emberae Morón, 2015
Phyllophaga emburyi Saylor, 1938
Phyllophaga eniba Saylor, 1943
Phyllophaga ephilida (Say, 1825)
Phyllophaga epigaea (Wickham, 1903)
Phyllophaga epulara Sanderson, 1958
Phyllophaga erota Saylor, 1942
Phyllophaga errans (LeConte, 1859)
Phyllophaga espina Sanderson, 1951
Phyllophaga esquinada Sanderson, 1951
Phyllophaga etundutejiana Morón & Hernández-Cruz, 2013
Phyllophaga euryaspis (Bates, 1887)
Phyllophaga exorata (Horn, 1887)
Phyllophaga expansa Chapin, 1932
Phyllophaga explanicollis Chapin, 1935
Phyllophaga falcata Smith & Paulsen, 2015
Phyllophaga falta
Phyllophaga farcta (LeConte, 1856)
Phyllophaga favosa Saylor, 1941
Phyllophaga ferrugata (Moser, 1918)
Phyllophaga ferruginea (Moser, 1918)
Phyllophaga ferupilis Saylor, 1941
Phyllophaga fervida (Fabricius, 1775)
Phyllophaga fesina Saylor, 1948
Phyllophaga fimbriata Cahpin, 1932
Phyllophaga fissilabris (Bates, 1887)
Phyllophaga flavidopilosa (Moser, 1921)
Phyllophaga floridana Robinson, 1938
Phyllophaga forbesi Glasgow, 1916
Phyllophaga forcipata (Burmeister, 1855)
Phyllophaga forsteri (Burmeister, 1855)
Phyllophaga fossoria Sanderson, 1951
Phyllophaga foveicollis (Bates, 1887)
Phyllophaga foxii Davis, 1920
Phyllophaga fragilipennis (Blanchard, 1850)
Phyllophaga franciscana García-Vidál, 1984
Phyllophaga fraterna Harris, 1841
Phyllophaga freeborni Saylor, 1937
Phyllophaga fucata (Horn, 1887)
Phyllophaga fulvipennis (Blanchard, 1850)
Phyllophaga fulviventris (Moser, 1918)
Phyllophaga fusca (Frölich, 1792)
Phyllophaga fuscipennis (Moser, 1918)
Phyllophaga futilis (LeConte, 1850)
Phyllophaga gaigei Sanderson, 1948
Phyllophaga galeanae Saylor, 1943
Phyllophaga garrota Sanderson, 1951
Phyllophaga gastonguzmani Morón, 2016
Phyllophaga gaumeri Bates, 1890
Phyllophaga gentryi (Saylor, 1936)
Phyllophaga georgiana (Horn, 1885)
Phyllophaga ghentata (Saylor, 1937)
Phyllophaga ginigra Saylor, 1940
Phyllophaga glaberrima (Blanchard, 1850)
Phyllophaga glabricula (LeConte, 1856)
Phyllophaga gloriae Morón, 2013
Phyllophaga godwini Morón & Riley, 2005
Phyllophaga gracilis (Burmeister, 1855)
Phyllophaga gracillima (Dalla Torre, 1912)
Phyllophaga gramma Morón & Woodruff, 2014
Phyllophaga grancha Saylor, 1948
Phyllophaga gravidula (Moser, 1921)
Phyllophaga grossepunctata (Moser, 1918)
Phyllophaga guanacasteca Morón & Solis, 2000
Phyllophaga guanahacabibensis Serrano & Morón, 2017
Phyllophaga guanicana Smyth, 1917
Phyllophaga guapilana Saylor, 1935
Phyllophaga guapiles Saylor, 1941
Phyllophaga guapilesea Saylor, 1941
Phyllophaga guapoloides Morón & Solis, 2001
Phyllophaga guatemala Saylor, 1940
Phyllophaga guerrocans Saylor, 1938
Phyllophaga haagi Saylor, 1943
Phyllophaga haitiensis Woodruff, 2005
Phyllophaga hamata (Horn, 1887)
Phyllophaga hardyi García-Vidál, 1984
Phyllophaga hemilissa (Bates, 1887)
Phyllophaga hernandezi García-Vidál, 1984
Phyllophaga herrerana Morón, 2017
Phyllophaga heteroclita (Burmeister, 1855)
Phyllophaga heteronycha (Bates, 1887)
Phyllophaga hidalgoana Saylor, 1943
Phyllophaga hintonella Saylor, 1941
Phyllophaga hintoni Saylor, 1935
Phyllophaga hirsuta (Knoch, 1801)
Phyllophaga hirticollis (Moser, 1921)
Phyllophaga hirticula (Knoch, 1801)
Phyllophaga hirtifrons (Moser, 1921)
Phyllophaga hirtiventris (Horn, 1887)
Phyllophaga hoegei (Bates, 1887)
Phyllophaga hoegella Saylor, 1943
Phyllophaga hoffmanitae Morón, 1996
Phyllophaga hogardi (Blanchard, 1850)
Phyllophaga holguinensis Smith & Paulsen, 2015
Phyllophaga hondura Saylor, 1943
Phyllophaga hondurasana (Moser, 1921)
Phyllophaga hoogstraali Saylor, 1943
Phyllophaga hornii (Smith, 1889)
Phyllophaga howdenryi Morón & Maes, 2014
Phyllophaga huaiandehuana Morón & Hernández-Cruz, 2013
Phyllophaga hubbelli Cartwright, 1946
Phyllophaga huiteaca Morón, 2006
Phyllophaga humboldtiana Morón, 2003
Phyllophaga idonea Sanderson, 1948
Phyllophaga ignava (Horn, 1887)
Phyllophaga ilicis (Knoch, 1801)
Phyllophaga imeldae Morón, 2017
Phyllophaga imitans Frey, 1975
Phyllophaga imitatrix Chapin, 1932
Phyllophaga impar Davis, 1920
Phyllophaga implicita (Horn, 1887)
Phyllophaga impressipyga Frey, 1975
Phyllophaga imprima Sanderson, 1951
Phyllophaga incuria Sanderson, 1942
Phyllophaga independentista García-Vidál, 1984
Phyllophaga inepta (Horn, 1887)
Phyllophaga infidelis (Horn, 1887)
Phyllophaga inflativentris (Moser, 1918)
Phyllophaga inflexa Barrett, 1935
Phyllophaga insulaepinora García-Vidál, 1978
Phyllophaga insulana (Moser, 1918)
Phyllophaga insularis
Phyllophaga integra (Say, 1835)
Phyllophaga integriceps (Moser, 1918)
Phyllophaga inversa (Horn, 1887)
Phyllophaga invisa Riley & Wolfe, 1995
Phyllophaga irazuana Saylor, 1936
Phyllophaga isabellae Morón & Rivera, 2001
Phyllophaga itsmica Morón, 2000
Phyllophaga iviei Chalumeau, 1985
Phyllophaga izabalana Morón, 2003
Phyllophaga izucarana Morón & Aragon, 1997
Phyllophaga jalisciensis Morón, Rivera & López, 2001
Phyllophaga jamaicana (Moser, 1918)
Phyllophaga janzeniana Morón & Solis, 2000
Phyllophaga jaragua Woodruff, 2005
Phyllophaga jaronua Chapin, 1932
Phyllophaga javepacuana Morón, 2003
Phyllophaga jeanmathieui Morón & Woodruff, 2014
Phyllophaga jimenezi Woodruff & Sanderson, 2005
Phyllophaga jorgevaldezi Hernández-Cruz, Morón & Sánchez-García, 2015
Phyllophaga josepalaciosi Morón & Nogueira, 2012
Phyllophaga jovelana Morón & Cano, 2000
Phyllophaga joyana Morón, Rivera & López, 2001
Phyllophaga jumberea Saylor, 1942
Phyllophaga juquilana Morón, 2000
Phyllophaga karlsioei (Linell, 1897)
Phyllophaga kayaumariana Morón & Nogueira, 2014
Phyllophaga kenscoffi Wolcott, 1928
Phyllophaga kentuckiana Ritcher, 1937
Phyllophaga knausi (Schaeffer, 1907)
Phyllophaga knochi (Gyllenhal, 1817)
Phyllophaga kohlmanniana Morón & Solis, 2000
Phyllophaga kulzeri Frey, 1975
Phyllophaga kuntzeni (Moser, 1921)
Phyllophaga laboriosa García-Vidál, 1988
Phyllophaga labrata Chapin, 1932
Phyllophaga lacroixi Paulian, 1947
Phyllophaga laevigata (Blanchard, 1850)
Phyllophaga laminata (Moser, 1921)
Phyllophaga lanata (Blanchard, 1850)
Phyllophaga lanceolata (Say, 1824)
Phyllophaga lanepta (Saylor, 1937)
Phyllophaga larimar Woodruff, 2005
Phyllophaga latefissa (Moser, 1918)
Phyllophaga latens (Arrow, 1900)
Phyllophaga latidens (Schaeffer, 1906)
Phyllophaga latifrons (LeConte, 1856)
Phyllophaga latiungula Wolcott, 1928
Phyllophaga lebasii (Blanchard, 1850)
Phyllophaga leonilae Morón, 1995
Phyllophaga leonina (Bates, 1887)
Phyllophaga leporina (Erichson, 1848)
Phyllophaga leprieuri (Blanchard, 1850)
Phyllophaga leptospica Sanderson, 1951
Phyllophaga lineata (Bates, 1887)
Phyllophaga lineatoides Morón, 2000
Phyllophaga linharesensis Frey, 1975
Phyllophaga lobata (Fall, 1908)
Phyllophaga lodingi Sanderson, 1939
Phyllophaga longicornis (Blanchard, 1850)
Phyllophaga longifoliata (Moser, 1921)
Phyllophaga longispina (Smith, 1889)
Phyllophaga longitarsa (Say, 1824)
Phyllophaga lorencita Morón & Solis, 2001
Phyllophaga lota Luginbill, 1928
Phyllophaga loxichana Morón, 2017
Phyllophaga luctuosa (Horn, 1887)
Phyllophaga lulaana Morón, 2000
Phyllophaga luridipennis (Moser, 1918)
Phyllophaga macasana (Moser, 1921)
Phyllophaga macrocera
Phyllophaga macrophylla (Bates, 1887)
Phyllophaga maculicollis (LeConte, 1863)
Phyllophaga maestrensis García-Vidál, 1984
Phyllophaga magnicornis (Moser, 1921)
Phyllophaga mali Wolcott, 1928
Phyllophaga manantleca Morón, Rivera & López, 2001
Phyllophaga manaosana (Moser, 1924)
Phyllophaga manchesterea Saylor, 1940
Phyllophaga mandevillea Saylor, 1940
Phyllophaga manibota Smith & Paulsen, 2015
Phyllophaga marcano Woodruff, 2005
Phyllophaga marcapatana (Moser, 1918)
Phyllophaga marginalis (LeConte, 1856)
Phyllophaga mariaegalante Chalumeau & Gruner, 1976
Phyllophaga mariana Fall, 1929
Phyllophaga marina García-Vidál, 1987
Phyllophaga martiana Saylor, 1943
Phyllophaga martinezi Frey, 1975
Phyllophaga martinezpalaciosi Morón, 1988
Phyllophaga matacapana Morón, 2003
Phyllophaga mella Sanderson, 1951
Phyllophaga menetriesi (Blanchard, 1850)
Phyllophaga mentalis Saylor, 1941
Phyllophaga mesophylla Morón & Rivera, 1992
Phyllophaga mexicana (Blanchard, 1850)
Phyllophaga micans (Knoch, 1801)
Phyllophaga microcerus (Arrow, 1933)
Phyllophaga microchaeta (Moser, 1924)
Phyllophaga microphylla (Moser, 1918)
Polyphylla micros (Bates, 1888)
Phyllophaga microsoma Chapin, 1932
Phyllophaga migratoria García-Vidál, 1984
Phyllophaga minutissima Wolcott, 1928
Phyllophaga mitlana Saylor, 1943
Phyllophaga moei Saylor, 1938
Phyllophaga molopia (Bates, 1887)
Phyllophaga monana (Moser, 1921)
Phyllophaga monrosi Frey, 1975
Phyllophaga montserratensis (Arrow, 1920)
Phyllophaga morganella Saylor, 1942
Phyllophaga morgani Saylor, 1938
Phyllophaga moserella Morón, 2016
Phyllophaga moseri Frey, 1965
Phyllophaga munizi Morón, 2008
Phyllophaga murina (Burmeister, 1855)
Phyllophaga muwieriana Morón & Nogueira, 2014
Phyllophaga nahui Morón & Solis, 2004
Phyllophaga nandalumia Morón & Riley, 2005
Phyllophaga naranjina Morón & Solis, 2001
Phyllophaga nasalis Chapin, 1935
Phyllophaga navassa Woodruff & Steiner, 2011
Phyllophaga navidad Morón & Woodruff, 2008
Phyllophaga nebulosa Polihronakis, 2007
Phyllophaga necaxa Saylor, 1943
Phyllophaga neglecta (Blanchard, 1850)
Phyllophaga nepida Saylor, 1941
Phyllophaga nevermannea Saylor, 1941
Phyllophaga nevermanni Saylor, 1935
Phyllophaga nevomeana Morón, 2006
Phyllophaga nigerrima (Bates, 1887)
Phyllophaga nigrita (Moser, 1918)
Phyllophaga nisuens (Saylor, 1937)
Phyllophaga nitida (LeConte, 1856)
Phyllophaga nitidicauda (Arrow, 1913)
Phyllophaga nitidicollis (Blanchard, 1850)
Phyllophaga nitididorsis Frey, 1975
Phyllophaga nosa Blackwelder, 1944
Phyllophaga nubipennis (Bates, 1887)
Phyllophaga nuda (Moser, 1918)
Phyllophaga nudipennis Frey, 1975
Phyllophaga nunezi Woodruff, 2005
Phyllophaga oaxaca Saylor, 1940
Phyllophaga oaxena Saylor, 1940
Phyllophaga obscura (Brenske, 1892)
Phyllophaga obsoleta Blanchard, 1850
Phyllophaga ocozocuana Morón, 2003
Phyllophaga odomi Saylor, 1943
Phyllophaga ohausi (Moser, 1921)
Phyllophaga okeechobea Robinson, 1948
Phyllophaga omani Sanderson, 1937
Phyllophaga omiltemia Bates, 1890
Phyllophaga onchophora Chapin, 1932
Phyllophaga onita Saylor, 1941
Phyllophaga onoreana Morón, 2003
Phyllophaga opaca (Moser, 1918)
Phyllophaga orosina (Moser, 1918)
Phyllophaga ortizi Woodruff, 2005
Phyllophaga ovalis Cartwright, 1939
Phyllophaga pachuca Saylor, 1943
Phyllophaga pachypyga (Burmeister, 1855)
Phyllophaga pallida (Horn, 1885)
Phyllophaga pallidicornis (Moser, 1921)
Phyllophaga pameana Morón, 2000
Phyllophaga panamana Chapin, 1935
Phyllophaga panamensis (Moser, 1921)
Phyllophaga panicula Sanderson, 1951
Phyllophaga panorpa Sanderson, 1950
Phyllophaga papaloana Morón, 2001
Phyllophaga paraguayana (Moser, 1921)
Phyllophaga paraguayensis (Moser, 1918)
Phyllophaga parallela (Blanchard, 1850)
Phyllophaga parcesetifera Chapin, 1935
Phyllophaga parilis (Bates, 1888)
Phyllophaga parumpunctata (Bates, 1887)
Phyllophaga parvicornis (Moser, 1924)
Phyllophaga parvidens (LeConte, 1856)
Phyllophaga parvisetis (Bates, 1887)
Phyllophaga pastassana (Moser, 1924)
Phyllophaga paternoi (Glasgow, 1925)
Phyllophaga patruelis (Chevrolat, 1865)
Phyllophaga patrueloides Paulian, 1947
Phyllophaga pauli Serrano & Morón, 2017
Phyllophaga pauliani Chalumeau & Gruner, 1976
Phyllophaga pauloensis Frey, 1975
Phyllophaga pearliae Davis, 1920
Phyllophaga pectoralis (Blanchard, 1850)
Phyllophaga pedernales Woodruff, 2005
Phyllophaga penaella Frey, 1975
Phyllophaga peninsulana (Moser, 1918)
Phyllophaga pentaphylla (Bates, 1887)
Phyllophaga perfidia Smith & Paulsen, 2015
Phyllophaga perlonga Davis, 1920
Phyllophaga permagna (Moser, 1918)
Phyllophaga persimilis Chapin, 1935
Phyllophaga peruana (Moser, 1918)
Phyllophaga picadoana Morón & Solis, 2000
Phyllophaga picea (Blanchard, 1850)
Phyllophaga piceola (Bates, 1887)
Phyllophaga picipennis (Moser, 1918)
Phyllophaga piligera (Moser, 1918)
Phyllophaga piliventris (Moser, 1918)
Phyllophaga pilositarsis Blackwelder, 1944
Phyllophaga pilosula (Moser, 1918)
Phyllophaga pilotoensis García-Vidál, 1987
Phyllophaga pilula (Moser, 1921)
Phyllophaga plaei (Blanchard, 1850)
Phyllophaga plairi (Saylor, 1937)
Phyllophaga platti Saylor, 1935
Phyllophaga platyrhina (Bates, 1887)
Phyllophaga pleroma Reinhard, 1940
Phyllophaga poculifer (Bates, 1887)
Phyllophaga pokornyiana Morón, 1995
Phyllophaga polita (Harold, 1869)
Phyllophaga polyphylla (Bates, 1887)
Phyllophaga postrema (Horn, 1887)
Phyllophaga poterillosa Saylor, 1942
Phyllophaga potosisalta Morón & Woodruff, 2008
Phyllophaga potrerillo García-Vidál, 1987
Phyllophaga praetermissa (Horn, 1887)
Phyllophaga probaporra Sanderson, 1951
Phyllophaga profunda (Blanchard, 1850)
Phyllophaga prolixa (Bates, 1887)
Phyllophaga propinqua (Moser, 1918)
Phyllophaga pruinipennis (Moser, 1918)
Phyllophaga pruinosa (Blanchard, 1850)
Phyllophaga prunina (LeConte, 1856)
Phyllophaga prununculina (Burmeister, 1855)
Phyllophaga pseudoatra Morón, 2003
Phyllophaga pseudocalcaris Saylor, 1940
Phyllophaga pseudocarus Morón, 1999
Phyllophaga pseudofloridana Woodruff & Beck, 1989
Phyllophaga pseudomicans Chapin, 1935
Phyllophaga psiloptera Sanderson, 1939
Phyllophaga puberea (Mannerheim, 1828)
Phyllophaga puberula (Jacquelin Du Val, 1856)
Phyllophaga pubescens (Burmeister, 1855)
Phyllophaga pubicauda (Bates, 1887)
Phyllophaga pubicollis (Blanchard, 1850)
Phyllophaga pudorosa Reinhard, 1939
Phyllophaga punctipennis (Blanchard, 1850)
Phyllophaga punctulata (Blanchard, 1850)
Phyllophaga punctuliceps (Bates, 1887)
Phyllophaga punctulicollis (Bates, 1887)
Phyllophaga puntarenosa Morón & Solis, 2001
Phyllophaga pusillidens Fall, 1937
Phyllophaga quadriphylla Saylor, 1943
Phyllophaga quercus (Knoch, 1801)
Phyllophaga quetzala Morón, 2001
Phyllophaga quetzaloides Morón, 2001
Phyllophaga quetzalpoa Morón, 2015
Phyllophaga quiana Morón, 2003
Phyllophaga quituana Morón, 2015
Phyllophaga rafaelamothei Morón, 2016
Phyllophaga rangelana Chapin, 1935
Phyllophaga ravida (Blanchard, 1850)
Phyllophaga rawlinsi Woodruff, 2005
Phyllophaga raydoma Saylor, 1943
Phyllophaga recorta Sanderson, 1951
Phyllophaga reevesi Saylor, 1939
Phyllophaga regiomontana Morón, 2001
Phyllophaga renodis Reinhard, 1939
Phyllophaga reticulata Frey, 1975
Phyllophaga reventazona Saylor, 1936
Phyllophaga rex Woodruff & Sanderson, 2005
Phyllophaga reyescastilloi Morón, 2013
Phyllophaga reyesiana Morón, 1992
Phyllophaga riviera Reinhard. 1950
Phyllophaga roberterroni Morón, 2017
Phyllophaga rolbakeri Saylor, 1940
Phyllophaga rolstoni Riley & Wolfe, 1995
Phyllophaga romana (Saylor, 1946)
Phyllophaga rorida (Burmeister, 1855)
Phyllophaga rorulenta (Burmeister, 1855)
Phyllophaga roscida (Burmeister, 1855)
Phyllophaga rossi Saylor, 1939
Phyllophaga rostripyga (Bates, 1890)
Phyllophaga rubiginosa (LeConte, 1856)
Phyllophaga rubricosa Reinhard, 1939
Phyllophaga ruficollis (Moser, 1915)
Phyllophaga rufipes (Moser, 1924)
Phyllophaga rufithorax (Moser, 1921)
Phyllophaga rufiventris (Kirsch, 1885)
Phyllophaga rufotestacea (Moser, 1918)
Phyllophaga rugicollis (Bates, 1887)
Phyllophaga rugithorax Saylor, 1938
Phyllophaga rugosa (Melsheimer, 1845)
Phyllophaga rugulosa (Blanchard, 1850)
Phyllophaga rustica Woodruff, 2005
Phyllophaga rzedowskiana Aragon & Morón, 2003
Phyllophaga sabanalamarensis Serrano & Morón, 2017
Phyllophaga sacoma Reinhard, 1939
Phyllophaga saginata (Mannerheim, 1828)
Phyllophaga sahagundinoi Morón, 2015
Phyllophaga saltana Frey, 1975
Phyllophaga sanbarthensis Chalumeau & Gruner, 1976
Phyllophaga sanctipauli (Blanchard, 1850)
Phyllophaga sandersoni García-Vidál, 1988
Phyllophaga sandersoniana Morón, 1992
Phyllophaga sandersoniella Chalumeau & Gruner, 1976
Phyllophaga santachloe Woodruff, 2005
Phyllophaga santaclarae Chapin, 1932
Phyllophaga santiagozai Morón, 2011
Phyllophaga santiaguensis García-Vidál, 1987
Phyllophaga sarlacca Smith & Paulsen, 2015
Phyllophaga saylori Sanderson, 1965
Phyllophaga sayloriana Morón, Rivera & López, 2001
Phyllophaga scabrifrons (Bates, 1887)
Phyllophaga scaramuzzai García-Vidál, 1988
Phyllophaga schaefferi Saylor, 1937
Phyllophaga schenklingi (Moser, 1918)
Phyllophaga schizorhina (Bates, 1887)
Phyllophaga schizorhinoides Morón, 2003
Phyllophaga schneblei Frey, 1975
Phyllophaga schusteriana Cano & Morón, 2002
Phyllophaga schwarzi Chapin, 1932
Phyllophaga scissa (Bates, 1887)
Phyllophaga scitula (Horn, 1887)
Phyllophaga secessicola Smith & Paulsen, 2015
Phyllophaga sechi Cano, 2020
Phyllophaga segregans (Bates, 1887)
Phyllophaga senicula (Bates, 1887)
Phyllophaga sequoiana Saylor, 1936
Phyllophaga sericata (Erichson, 1848)
Phyllophaga serrana Morón & Cano, 2000
Phyllophaga serratipes Morón & Cano, 2000
Phyllophaga sibonyensis García-Vidál, 1988
Phyllophaga signaticollis (Burmeister, 1855)
Phyllophaga sinaloana Saylor, 1935
Phyllophaga sinicollis Saylor, 1943
Phyllophaga sinuaticeps (Moser, 1921)
Phyllophaga skelleyi Woodruff & Beck, 1989
Phyllophaga smithi (Arrow, 1912)
Phyllophaga sodalis Reinhard, 1940
Phyllophaga solavegana Morón, 2000
Phyllophaga solisiana Morón, 2003
Phyllophaga sonora Saylor, 1939
Phyllophaga soror Davis, 1920
Phyllophaga spaethi (Nonfried, 1891)
Phyllophaga speculifera (Chevrolat, 1865)
Phyllophaga spinicola García-Vidál, 1984
Phyllophaga spinifemora Saylor, 1940
Phyllophaga spinitarsis (Moser, 1921)
Phyllophaga spreta (Horn, 1887)
Phyllophaga squamifera Frey, 1976
Phyllophaga squamipilosa Saylor, 1936
Phyllophaga stehlei Chalumeau, 1985
Phyllophaga stipitalis (Blanchard, 1850)
Phyllophaga stohleri Saylor, 1938
Phyllophaga straminea (Bates, 1887)
Phyllophaga sturmi (Bates, 1887)
Phyllophaga stzotzilana Morón, 2001
Phyllophaga submetallica (Bates, 1887)
Phyllophaga submucida (LeConte, 1856)
Phyllophaga subnitida (Moser, 1918)
Phyllophaga subopaca (Moser, 1918)
Phyllophaga subpruinosa (Casey, 1884)
Phyllophaga subrugosa (Moser, 1924)
Phyllophaga subtonsa (LeConte, 1856)
Phyllophaga suttonana Reinhard, 1939
Phyllophaga suturalis (Chevrolat, 1865)
Phyllophaga sylvatica Sanderson, 1942
Phyllophaga tajimaroana Morón, 2000
Phyllophaga talamancana Morón & Solis, 2001
Phyllophaga tamatziana Morón & Nogueira, 2014
Phyllophaga tancitara Saylor, 1943
Phyllophaga tapantina Morón & Solis, 2001
Phyllophaga taxodii Langston, 1924
Phyllophaga tecta Cartwright, 1944
Phyllophaga tegenara Saylor, 1935
Phyllophaga tegulicollis Saylor, 1934
Phyllophaga tejulpicas Saylor, 1943
Phyllophaga tejupana Morón, 2000
Phyllophaga temascalis Saylor, 1941
Phyllophaga temascaltepeca Saylor, 1934
Phyllophaga temaxa Saylor, 1940
Phyllophaga temora Saylor, 1943
Phyllophaga tenuipilis (Bates, 1887)
Phyllophaga teosinteophaga Morón & Rivera, 1992
Phyllophaga tepanana Saylor, 1938
Phyllophaga tepequetzala Morón, 2015
Phyllophaga terezinae Frey, 1965
Phyllophaga tesorito Vallejo & Wolff, 2013
Phyllophaga tetracera Chapin, 1932
Phyllophaga tetraphylla (Moser, 1918)
Phyllophaga thoracica (Burmeister, 1855)
Phyllophaga tilarana Morón & Sólis, 2001
Phyllophaga tinbitipus Morón, 2013
Phyllophaga tlamana Morón, 2016
Phyllophaga tlanchinolensis Morón, 2000
Phyllophaga tlilloa Morón, 2018
Phyllophaga tojolabala Morón, 1999
Phyllophaga toni Woodruff, 2005
Phyllophaga torta (LeConte, 1856)
Phyllophaga totonis Saylor, 1941
Phyllophaga totoreana Morón, 2006
Phyllophaga trichoderma (Moser, 1921)
Phyllophaga trichodes (Bates, 1887)
Phyllophaga tridens (Bates, 1887)
Phyllophaga tridilonycha Saylor, 1943
Phyllophaga trinitariensis García-Vidál, 1987
Phyllophaga triticophaga Morón & Salvadori, 1998
Phyllophaga trochanter Saylor, 1940
Phyllophaga tsajumiana Morón, 2001
Phyllophaga tuberculifrons (Chevrolat, 1865)
Phyllophaga tumulosa Bates, 1888
Phyllophaga tusa (Horn, 1887)
Phyllophaga tuxtleca Morón, 2003
Phyllophaga tzeltala Morón, 2013
Phyllophaga ueiacayoca Morón, 1991
Phyllophaga ulkei (Smith, 1889)
Phyllophaga umbrosa (Erichson, 1847)
Phyllophaga uniformis (Blanchard, 1850)
Phyllophaga uruguayana (Saylor, 1935)
Phyllophaga valeriana Saylor, 1934
Phyllophaga vallendensis Morón, 2013
Phyllophaga vandinei Smyth, 1917
Phyllophaga vandykei Saylor, 1935
Phyllophaga varohiana Morón, 2006
Phyllophaga vazquezae Morón, 1995
Phyllophaga vehemens (Horn, 1887)
Phyllophaga velezangeli Morón & Neita-Moreno, 2014
Phyllophaga vermiculata Chapin, 1932
Phyllophaga verruciventris (Moser, 1918)
Phyllophaga vestita (Moser, 1918)
Phyllophaga vicina (Moser, 1918)
Phyllophaga vilifrons (LeConte, 1856)
Phyllophaga villaclarensis García-Vidál, 1987
Phyllophaga villardoi Morón & Ordóñez-Résendiz, 2015
Phyllophaga violetae Morón & Aragon-García, 2012
Phyllophaga vulpes (Arrow, 1913)
Phyllophaga wittkugeli (Nonfried, 1891)
Phyllophaga wolcotti Saylor, 1940
Phyllophaga xalacoatepecana Morón, 2015
Phyllophaga xanthe (Bates, 1887)
Phyllophaga xanthocoma (Bates, 1887)
Phyllophaga xerophila Saylor, 1939
Phyllophaga xkumuka Morón, 1999
Phyllophaga yautepecana Morón, 2018
Phyllophaga yaxbitana Morón, 2000
Phyllophaga yemasseei Cartwright, 1944
Phyllophaga yoloxana Morón, 2003
Phyllophaga youngi Cartwright, 1935
Phyllophaga yucana Saylor, 1937
Phyllophaga yunqueana Chapin, 1935
Phyllophaga zacaquetzala Morón, 2015
Phyllophaga zaragozana Morón, 2003
Phyllophaga zarcoana Morón, 2003
Phyllophaga zavalana Reinhard, 1946
Phyllophaga zayasi García-Vidál, 1978
Phyllophaga zeteki Saylor, 1942
Phyllophaga zunimarioi Morón & Maes, 2014

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Generic Guide to New World Scarab Beetles - Phyllophaga [1]

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