Lonchopteridae

Family of flies

Lonchopteridae
Lonchoptera lutea
Scientific classification Edit this classification
Domain: Eukaryota
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Order: Diptera
Superfamily: Phoroidea
Family: Lonchopteridae
Macquart, 1823[1]
Diversity
6 genera
Synonyms
  • Lonchopteroidea
  • Musidoridae Kertész, 1909[2]
wing venation (male)

The Lonchopteridae (spear-winged flies or pointed-wing flies) are a family of small (2–5 mm), slender, yellow to brownish-black Diptera, occurring all over the world. Their common name refers to their pointed wings, which have a distinct venation. Many are parthenogenic; males are very rare, however, at least in North American species, and have a somewhat different venation than do the females.[3][4]

Spear-winged flies are common in moist, shady, grassy areas, where the larvae are found within decaying vegetation. One species, Lonchoptera bifurcata, is cosmopolitan in distribution, and may have been transported via shipments of vegetables.[3][4]

Description

The Lonchopteridae are minute, slender flies with long wings which are pointed at the apex. The head is rounded, with the outer vertical bristles, inner vertical bristles, ocellar bristles, interfrontal bristles, and bristles along the margin of the broad mouth very well developed. The mesonotum and scutellum and legs have well developed bristles. The radial vein R has three branches (R1, R2+3, R4+5). The median vein M is furcate (M1, M2). The anal vein A merges with the cubital vein Cu (female) or terminates freely (male).

Systematics

They are usually placed in the superfamily of flat-footed flies and allies (Platypezoidea). If the Platypezoidea are restricted to the flat-footed flies sensu stricto, the spear-winged flies are united with the Ironomyiidae and the coffin and scuttle flies (Phoridae) as Phoroidea. More rarely, they are treated as monotypic superfamily Lonchopteroidea.

Four[5] living genera are in this family, encompassing some 50 described species all together:[6]

  • Homolonchoptera Yang, 1998[7]
  • Lonchoptera Meigen, 1803[8]
  • Neolonchoptera Vaillant, 1989[9]
  • Spilolonchoptera Yang, 1998[7]

Two fossil genera of spear-winged flies have been described:[6]

  • Lonchopterites Grimaldi & Cumming, 1999[10]
  • Lonchopteromorpha Grimaldi & Cumming, 1999[10]

Species

  • West Palaearctic including Russia
  • Australasian/Oceanian
  • Nearctic
  • Japan
  • World list

References

  1. ^ Macquart, P.J.M. (1835). Histoire Naturelle des insectes. Diptères. Tome deuxieme. Paris: Roret. pp. 703 or 710 pp., 12 pls. Retrieved 27 January 2021.
  2. ^ Kertész, Kálmán (1909). Catalogus Dipterorum hucusque Descriptorum, Volume VI: Empididae, Dolichopodidae, Musidoridae. Budapest: Museum National Hungaricum. pp. 1–362. Retrieved 17 August 2016.
  3. ^ a b Smith, K.G.V. (1969): Handbook for the Identification of British Insects 10(2ai: Diptera Lonchopteridae): 1–9.
  4. ^ a b Borror, D.J.; Triplehorn, C.A. & Johnson, N.A. (1989): An Introduction to the Study of Insects (6th edition). Saunders College Publishing.
  5. ^ Tree of Life Web Project (2007). "Lonchopteridae - Spear-winged flies". Tree of Life Web Project.
  6. ^ a b Tree of Life Web Project (ToL) (2007): Lonchopteridae. Version of 2007-NOV-29. Retrieved 2009-APR-07.
  7. ^ a b Yang, C.K. (1998). Lonchopteridae, In: Xue, W.Q. & Chao, C.M. (eds.), Flies of China. Vol. 1. Shenyang: Liaoning Science and Technology Press. pp. 49–59.
  8. ^ Meigen, J. W. (1803). "Versuch einer neuen Gattungs-Eintheilung der europaischen zweiflugligen Insekten". Mag. Insektenkd. 2: 259–281.
  9. ^ Vaillant, François (1989). "Contribution à l'étude des diptères Lonchopteridae d'Europe et d'Afrique du nord". Bulletin de la Société Vaudoise des Sciences Naturelles. 79: 209–229.
  10. ^ a b Grimaldi, D.A.; Cumming, J.M. (1999). "Brachyceran Diptera in Cretaceous ambers and Mesozoic diversification of the Eremoneura". Bulletin of the American Museum of Natural History. 239: 1–124. hdl:2246/1583.
  • Lonchopteridae In Italian
  • Lonchopteridae page at the Bishop Museum, Honolulu
  • Family Lonchopteridae at EOL Image Gallery
  • Photograph of Lonchoptera furcata Fallén Should be L. bifurcata.
  • Photograph of Lonchoptera lutea Panzer
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Extant Diptera families
Suborder Nematocera
Axymyiomorpha
Culicomorpha
Culicoidea
  • Dixidae (meniscus midges)
  • Corethrellidae (frog-biting midges)
  • Chaoboridae (phantom midges)
  • Culicidae (mosquitoes)
Chironomoidea
  • Thaumaleidae (solitary midges)
  • Simuliidae (black flies)
  • Ceratopogonidae (biting midges)
  • Chironomidae (non-biting midges)
Blephariceromorpha
  • Blephariceridae (net-winged midges)
  • Deuterophlebiidae (mountain midges)
  • Nymphomyiidae
Bibionomorpha
Bibionoidea
  • Bibionidae (march flies, lovebugs)
Anisopodoidea
  • Anisopodidae (wood gnats)
Sciaroidea
(fungus gnats)
Perissommatomorpha
Psychodomorpha
Scatopsoidea
Psychodoidea
  • Psychodidae (moth flies)
Ptychopteromorpha
  • Ptychopteridae (phantom crane flies)
  • Tanyderidae (primitive crane flies)
Tipulomorpha
Trichoceroidea
  • Trichoceridae (winter crane flies)
Tipuloidea
(crane flies)
  • Cylindrotomidae (long-bodied crane flies)
  • Limoniidae (limoniid crane flies)
  • Pediciidae (hairy-eyed craneflies)
  • Tipulidae (large crane flies)
Suborder Brachycera
Asilomorpha
Asiloidea
Empidoidea
Nemestrinoidea
  • Acroceridae (small-headed flies)
  • Nemestrinidae (tangle-veined flies)
Muscomorpha
Aschiza
Platypezoidea
  • Ironomyiidae (ironic flies)
  • Lonchopteridae (spear-winged flies)
  • Opetiidae (flat-footed flies)
  • Phoridae (scuttle flies, coffin flies, humpbacked flies)
  • Platypezidae (flat-footed flies)
Syrphoidea
  • Pipunculidae (big-headed flies)
  • Syrphidae (hoverflies)
Schizophora
Acalyptratae
Conopoidea
  • Conopidae (thick-headed flies)
Tephritoidea
  • Pallopteridae (flutter flies)
  • Piophilidae (cheese flies)
  • Platystomatidae (signal flies)
  • Pyrgotidae
  • Richardiidae
  • Tephritidae (peacock flies)
  • Ulidiidae (picture-winged flies)
Nerioidea
  • Cypselosomatidae
  • Micropezidae (stilt-legged flies)
  • Neriidae (cactus flies, banana stalk flies)
Diopsoidea
Sciomyzoidea
Sphaeroceroidea
Lauxanioidea
Opomyzoidea
Ephydroidea
  • Camillidae
  • Curtonotidae (quasimodo flies)
  • Diastatidae (bog flies)
  • Drosophilidae (vinegar and fruit flies)
  • Ephydridae (shore flies)
  • Mormotomyiidae (frightful hairy fly)
Carnoidea
Lonchaeoidea
Calyptratae
Muscoidea
  • Anthomyiidae (cabbage flies)
  • Fanniidae (little house flies)
  • Muscidae (house flies, stable flies)
  • Scathophagidae (dung flies)
Oestroidea
Hippoboscoidea
  • Glossinidae (tsetse flies)
  • Hippoboscidae (louse flies)
  • Nycteribiidae (bat flies)
  • Streblidae (bat flies)
Stratiomyomorpha
Stratiomyoidea
  • Pantophthalmidae (timber flies)
  • Stratiomyidae (soldier flies)
  • Xylomyidae (wood soldier flies)
Tabanomorpha
Rhagionoidea
Tabanoidea
Vermileonomorpha
Vermileonoidea
Xylophagomorpha
Xylophagoidea
  • Xylophagidae (awl flies)
Taxon identifiers
Lonchopteridae